Arguably the most successful baseball coach in UCLA history, John Savage has reached a new milestone: 400 wins.

The 12th-year head coach earned that victory in dramatic fashion on Sunday night, as the No. 14 Bruins (6-5) pushed past USC, 5-3, in a 14-inning thriller. Senior outfielder Christoph Bono batted in UCLA’s go-ahead runs with a bases-loaded chopper in the top of the final frame at Dodger Stadium.

Savage now has a 27-14 record against the Trojans, more wins than he has against any other opponent. The two teams will meet again on May 13 to start a three-game series at Jackie Robinson Stadium.

The 51-year-old now is the fourth active Pac-12 coach to hit the 400-win threshold, joining Stanford’s Mark Marquess, Oregon State’s Pat Casey, and Cal’s David Esquer — and is the only one except Marquess to do it in fewer than 12 full seasons.

He remains the Bruins’ third-winningest coach behind Gary Adams (985) and Art Reichle (747), but holds a higher winning percentage (.592) and brought back the program’s only national title in 2013. Savage is under contract through 2025.